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This project brings together scholars in philosophy and biology to address the problem of integrating different types of explanations from scientific approaches to better understand life’s complexity. The specific focus is the question of how to bridge the divide between genetic and generic (physical) explanatory approaches in three domains of biological phenomena: development, evolutionary novelty, and evolvability. In order to facilitate new forms of explanatory integration, encourage progress on these questions, and generate epistemic models that assist ongoing inquiry, a core team of scientists and philosophers with relevant expertise will meet with a select group of internationally renowned experts for three intensive workshops with each year devoted to one domain.